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Dire wolf

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Dire wolf

The dire wolf (Aenocyon dirus ) is an extinct species of canine which was native to the Americas during the Late Pleistocene and Early Holocene epochs (125,000–10,000 years ago). The species was named in 1858, four years after the first specimen had been found. Two subspecies are proposed, Aenocyon dirus guildayi and Aenocyon dirus dirus, but this assignment has been recently considered questionable. The largest collection of its fossils has been obtained from the Rancho La Brea Tar Pits in Los Angeles. Dire wolf remains have been found across a broad range of habitats including plains, grasslands, and some forested mountain areas of North America and the arid savanna of South America. The sites range in elevation from sea level to 2,255 meters (7,400 ft). Dire wolf fossils have rarely been found north of 42°N latitude; there have been only five unconfirmed records above this latitude. This range restriction is thought to be due to temperature, prey, or habitat limitations imposed by proximity to the Laurentide and Cordilleran ice sheets that existed at the time. The dire wolf was about the same size as the largest modern forms of gray wolf (Canis lupus). A. d. guildayi weighed on average 60 kilograms (132 lb) and A. d. dirus was on average 68 kg (150 lb). Its skull and dentition matched those of C. lupus, but its teeth were larger with greater shearing ability, and its bite force at the canine tooth was stronger than any known Canis species. These characteristics are thought to be adaptations for preying on Late Pleistocene megaherbivores; in North America, its prey is suggested to have included western horses, dwarf pronghorn, flat-headed peccary, ground sloths, ancient bison, and camels. Dire wolves lived as recently as 10,000 years ago, according to dated remains. Its extinction occurred during the Quaternary extinction event, disappearing along with its main prey species; its reliance on megaherbivores has been proposed as the cause of its extinction, along with climatic change and competition with other species, or a combination of those factors.

Tables

Aenocyon dirus guildayi compared with the Yukon wolf by the mean length of limb bones in millimeters (inches) · Description
Humerus (upper front leg)
Humerus (upper front leg)
Limb variable
Humerus (upper front leg)
A. d. guildayi
218 mm (8.6 in)
Yukon wolf
237 mm (9.3 in)
A. d. dirus
240 mm (9.4 in)
Radius (lower front leg)
Radius (lower front leg)
Limb variable
Radius (lower front leg)
A. d. guildayi
209 mm (8.2 in)
Yukon wolf
232 mm (9.1 in)
A. d. dirus
240 mm (9.4 in)
Metacarpal (front foot)
Metacarpal (front foot)
Limb variable
Metacarpal (front foot)
A. d. guildayi
88 mm (3.4 in)
Yukon wolf
101 mm (4.0 in)
A. d. dirus
101 mm (4.0 in)
Femur (upper back leg)
Femur (upper back leg)
Limb variable
Femur (upper back leg)
A. d. guildayi
242 mm (9.5 in)
Yukon wolf
251 mm (9.9 in)
A. d. dirus
266 mm (10.5 in)
Tibia (lower back leg)
Tibia (lower back leg)
Limb variable
Tibia (lower back leg)
A. d. guildayi
232 mm (9.1 in)
Yukon wolf
258 mm (10.2 in)
A. d. dirus
255 mm (10.0 in)
Metatarsal (back foot)
Metatarsal (back foot)
Limb variable
Metatarsal (back foot)
A. d. guildayi
93 mm (3.7 in)
Yukon wolf
109 mm (4.3 in)
A. d. dirus
107 mm (4.2 in)
Limb variable
A. d. guildayi
Yukon wolf
A. d. dirus
Humerus (upper front leg)
218 mm (8.6 in)
237 mm (9.3 in)
240 mm (9.4 in)
Radius (lower front leg)
209 mm (8.2 in)
232 mm (9.1 in)
240 mm (9.4 in)
Metacarpal (front foot)
88 mm (3.4 in)
101 mm (4.0 in)
101 mm (4.0 in)
Femur (upper back leg)
242 mm (9.5 in)
251 mm (9.9 in)
266 mm (10.5 in)
Tibia (lower back leg)
232 mm (9.1 in)
258 mm (10.2 in)
255 mm (10.0 in)
Metatarsal (back foot)
93 mm (3.7 in)
109 mm (4.3 in)
107 mm (4.2 in)
Canis lupus and Aenocyon dirus compared by mean mandible tooth measurements (millimeters) · Adaptation › Dentition and bite force
m1 length
m1 length
Tooth variable
m1 length
lupus modern North American
28.2
lupus La Brea
28.9
lupus Beringia
29.6
dirus dirus Sangamonian era (125,000–75,000 YBP)
36.1
dirus dirus Late Wisconsin (50,000 YBP)
35.2
dirus guildayi (40,000–13,000 YBP)
33.3
m1 width
m1 width
Tooth variable
m1 width
lupus modern North American
10.7
lupus La Brea
11.3
lupus Beringia
11.1
dirus dirus Sangamonian era (125,000–75,000 YBP)
14.1
dirus dirus Late Wisconsin (50,000 YBP)
13.4
dirus guildayi (40,000–13,000 YBP)
13.3
m1 trigonid length
m1 trigonid length
Tooth variable
m1 trigonid length
lupus modern North American
19.6
lupus La Brea
21.9
lupus Beringia
20.9
dirus dirus Sangamonian era (125,000–75,000 YBP)
24.5
dirus dirus Late Wisconsin (50,000 YBP)
24.0
dirus guildayi (40,000–13,000 YBP)
24.4
p4 length
p4 length
Tooth variable
p4 length
lupus modern North American
15.4
lupus La Brea
16.6
lupus Beringia
16.5
dirus dirus Sangamonian era (125,000–75,000 YBP)
16.7
dirus dirus Late Wisconsin (50,000 YBP)
16.0
dirus guildayi (40,000–13,000 YBP)
19.9
p4 width
p4 width
Tooth variable
p4 width
lupus modern North American
-
lupus La Brea
-
lupus Beringia
-
dirus dirus Sangamonian era (125,000–75,000 YBP)
10.1
dirus dirus Late Wisconsin (50,000 YBP)
9.6
dirus guildayi (40,000–13,000 YBP)
10.3
p2 length
p2 length
Tooth variable
p2 length
lupus modern North American
-
lupus La Brea
-
lupus Beringia
-
dirus dirus Sangamonian era (125,000–75,000 YBP)
15.7
dirus dirus Late Wisconsin (50,000 YBP)
14.8
dirus guildayi (40,000–13,000 YBP)
15.7
p2 width
p2 width
Tooth variable
p2 width
lupus modern North American
-
lupus La Brea
-
lupus Beringia
-
dirus dirus Sangamonian era (125,000–75,000 YBP)
7.1
dirus dirus Late Wisconsin (50,000 YBP)
6.7
dirus guildayi (40,000–13,000 YBP)
7.4
Tooth variable
lupus modern North American
lupus La Brea
lupus Beringia
dirus dirus Sangamonian era (125,000–75,000 YBP)
dirus dirus Late Wisconsin (50,000 YBP)
dirus guildayi (40,000–13,000 YBP)
m1 length
28.2
28.9
29.6
36.1
35.2
33.3
m1 width
10.7
11.3
11.1
14.1
13.4
13.3
m1 trigonid length
19.6
21.9
20.9
24.5
24.0
24.4
p4 length
15.4
16.6
16.5
16.7
16.0
19.9
p4 width
-
-
-
10.1
9.6
10.3
p2 length
-
-
-
15.7
14.8
15.7
p2 width
-
-
-
7.1
6.7
7.4
Dire wolf information based on skull measurements[74]
Body size
Body size
Variable
Body size
28,000 YBP
largest
26,100 YBP
large
17,900 YBP
smallest
13,800 YBP
medium/small
Tooth breakage
Tooth breakage
Variable
Tooth breakage
28,000 YBP
high
26,100 YBP
low
17,900 YBP
high
13,800 YBP
low
Tooth wear
Tooth wear
Variable
Tooth wear
28,000 YBP
high
26,100 YBP
low
17,900 YBP
high
13,800 YBP
low
Snout shape
Snout shape
Variable
Snout shape
28,000 YBP
shortening, largest cranial base
26,100 YBP
average
17,900 YBP
shortest, largest cranial base
13,800 YBP
average
Tooth row shape
Tooth row shape
Variable
Tooth row shape
28,000 YBP
robust
26,100 YBP
17,900 YBP
13,800 YBP
gracile
DO event
DO event
Variable
DO event
28,000 YBP
number 3 or 4
26,100 YBP
none
17,900 YBP
imprecise data
13,800 YBP
imprecise data
Variable
28,000 YBP
26,100 YBP
17,900 YBP
13,800 YBP
Body size
largest
large
smallest
medium/small
Tooth breakage
high
low
high
low
Tooth wear
high
low
high
low
Snout shape
shortening, largest cranial base
average
shortest, largest cranial base
average
Tooth row shape
robust
gracile
DO event
number 3 or 4
none
imprecise data
imprecise data

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